Acer has always felt like an outcast in the car world. The beat-up green AMC Pacer joined forces with fellow “lemon” cars as henchmen for the devious Professor Z, whose clandestine mission is to wreak havoc at the highly visible World Grand Prix. Acer must hunt down the American and British secret agents who’ve stolen crucial information about Professor Z’s underhanded plot—his primary target just happens to be Mater, who’s been mistaken for a spy. Acer tries very hard to be a tough guy, but he’s over-eager compared to his no-nonsense accomplice Grem.
Grem is a dented, rusty orange AMC Gremlin. After years of being dismissed for his design, even being called a “lemon,” Grem has a big chip on his fender that has led him to the underworld of international espionage. As a henchman for a villainous boss Professor Z, Grem and his partner-in-crime Acer are trying to sabotage the World Grand Prix and the famous race cars competing in it. When Grem and Acer mistake Mater for an American agent with important top-secret information, the ruthless lemons set out on a round-the-globe chase to stop Mater from foiling their evil scheme. (Thanks for the tip Don)